Dogstorm and Cat Viper share their story of being on the journey with Oden and Roger but are interrupted by Nami who is concerned about the sea route. Meanwhile, Jack is plotting to bring down the Mokomo Dukedom again.
The plans are set. The Straw Hats divide and conquer, one group heading to save Sanji, the others to the Land of Wano. But in the final moments before their departure, the tremendous elephant island begins to quake.
After being aggressively attacked by Jack's fleet, the monstrous Zunesha comes to life and beckons to those on its back. But can the Straw Hats understand its cry? And realize in time what the elephant island needs from them?
With the skirmish with Jack now over, the Straw Hats move on to trying to bandage the great elephant's wounds. But just as the whole island puts the plan into action, unseen danger lurks on horizon.
The Straw Hats and the others split up into four teams, each on different missions. Some from the Mink Tribe insist on leaving with Luffy's team. Meanwhile, Usopp makes a weapon for Nami that seems to work like magic.
Rulers all across the world head for the quadrennial Levely, and Vivi gets in touch with her pirating past while at sea as she remembers her experiences with the Straw Hat Pirates.
Pedro discovers news regarding the Revolutionary Army attack caused by Blackbeard's pirates, and Luffy sees his father's face for the first time. Kaido is enraged by Doflamingo's capture, and he has captured a New Generation Pirate.
After spotting an island the Straw Hat's stop to get some food. however the island is home to a marine base and Luffy and the others must disguise themselves as marines if they want to get inside.
The jig is up as Luffy and the others make a daring dash to avoid the furious Navy. Can they escape without having to turn to fight - and after they've had a chance to raid the cabinets for food?
Making a mad dash for the Submerge with the food they stole, Luffy and the others have to get through some of the Navy's upstart soldiers. Grount, whose arm holds incredible power, summons his training to challenge Luffy.
The Straw Hats continue to suffer from blazing heat and a perilous food crisis. As the seas finally calm, fishing is their only hope. Meanwhile, Sanji arrives in Big Mom's territory and keeps his cool - until he sees his bride-to-be.
Luffy is in danger as the venomous fish poison courses through his body. With no antidotes on board, the Straw Hats must rely on a mysterious ship for help - a ship that bears a truly surprising crew.
On the edge of life and death, Luffy's only hope rests with Sanji's siblings. But even though they refuse to share an antidote, they may be able to save Luffy's life yet, giving him enough energy to plunge right back into trouble.
The Sanji Retrieval Team disembarks on Cacao Island. Luffy and Chopper eat an entire cafe and meet Charlotte Pudding, to whom Luffy accidentally reveals who he is. Big Mom oversees the preparation for the wedding Tea Party three days away.
The Straw Hats confront Pudding - the fiancee of Sanji. But when they challenge her for Sanji's heart, her answer shocks them. Meanwhile an ominous and mysterious message may spell doom for the Straw Hats.
The dangerous waters around the Whole Cake Island harbor terrifying creatures that pose a challenge for the Straw Hats. Meanwhile, Mama slips into a bout of her sickness and ravages her own land in search a specific pastry.
Big Mom continues her rampage and kills her son in a blind rage. Jinbe formally requests to leave her crew as he recalls his past experiences with Luffy. Sanji's Retrieval Team is attacked by fruit footballfish on which the team feasts.
Jinbei wants to leave Big Mom's crew but because Big Mom thinks of him as a powerful asset she wants him to lose something as well. Pedro tells the crew more about his past and the straw hats battle giant ants.
The straw hats arrive at the whole cake island. Pedro and Brook take the Shark Submerge 3 while Luffy, Carrot, Nami and Chopper look for Sanji. The crew see Sanji and go after him but they get distracted by sweets except for Nami.
Nami, Carrot and Chopper are chasing after Sanji, when doing so they meet a buried giant. The group is then attacked by a rabbit although the rabbit is not a Mink. Sanji fights one of his brothers.
Sanji beat his brother Yonji because of that Sanji's father, Judge Vinsmoke challenges him to a fight. A flashback reveals how Sanji's family mistreated and bullied him because they viewed him as weak.
News spread across the cake island about Jimbe in the newspaper. Pekom, captured by The Fire Tank Pirates was shot and falls off a cliff into sea full of sharks. Big Mom's plan for Caesar is revealed. Luffy is still stuck in the seducing woods.
Brûlée used the Mira Mira no Mi to trap Carrot in the Mirror-World. It is revealed why all flora and fauna are alive and how Big Mom taxes the Totto Land citizens. Big Mom now orders the attack of the Straw Hats and her ex-husband.
The identity of the enigmatic Pound, the man half-buried in the woods, is revealed, but his deep connection to Big Mom might not be enough to save him. Nami's past exploits, however, hold the key to keeping Mom's minions at bay.
A battle between Luffy and Cracker (one of the Sweet 3 Generals) begins. Nami is educated on some of the history of those who came to challenge Big Mom before.
Luffy continues to fight Cracker, and finds out that how troublesome his power can be. Brook and Pedro are on Cake Island and have to figure out how to move around the island without getting caught.
The wind starts to go out of Luffy's sails as he starts to fight an endless army of Biscuit Warriors. Meanwhile, the oldest two Vinsmoke brothers come home from a war to see their long-missing young brother Sanji.
Luffy cannot reach Cracker before Gear 4 runs out, and he has to hide while Biscuit Soldiers look for him. Judge threatens to order the destruction of the Baratie and have Zeff killed. Sanji remembers his history with Zeff.
Luffy needs just 10 minutes to recharge. Nami comes to rescue with the help of the Homies. Meanwhile, Sanji learns the terrible truth of Germa's power - and his father's dark history.
Nami challenges Cracker on her own. During the all-out fight, she loses her advantage and is close to defeat. Can Luffy bounce back in time? Meanwhile, Sanji's tragic childhood is made clear.
The terrible history of Sanji's family continue to unfold. From a rare moment of compassion he is able to escape, only to face his father's cruelty before setting out on his own.
Luffy filled with Cracker's biscuit, unlocks a new Gear Fourth attack that sends Cracker flying but it is not without consequences. Chopper and Carrot locate Sanji's reflection in one of Brûlée's mirror and try to communicate with him
Luffy refuses to fight Sanji, because of this Sanji beats Luffy. When Sanji is leaving Luffy tells him that he doesn't believe he meant what he said and that he'd be waiting for him.
Because Luffy beat Cracker Big Mom becomes enraged so she sends her enraged army to deal with him. The Vinsmokes arrive at the Whole Cake Chateau and have a feast with Big Mom.
Luffy faces Big Mom's enraged army. Nami and King Baum come back to help. In the Mirro-World, Randolph begins attacking Chopper and Carrot. Sanji agrees to marry pudding.
Brook manages to sneak into the Room of Treasure with the help of Pedro. While Sanji picks flowers for Pudding, she goes to the Prisoner Library to visit Luffy and Nami.
Pudding reveals a secret plan to prevent Sanji from marrying her, shocking Luffy and Nami and leaves saying goodbye. Brook defeats all the guards in treasure room. Tamago confronts Pedro in the third floor courtyard.
Chopper and carrot try to find a way out. Brook prepares to fight Big Mom. Pedro fights with baron and remembers how he fought him five years ago with Zepo.