Penguins discover a lost egg and decide hatching until the mother appears. When it opens unexpectedly, the new duck -Ducky- is trained as a penguin while a jealous King Julien wants to raise him to be his second in command.
Alex, Marty, Gloria and Melman are still fighting to get home to their beloved Big Apple. Their journey takes them through Europe where they find the perfect cover: a traveling circus, which they reinvent - Madagascar style.
Spoiled by their upbringing and unaware of what wildlife really is, four animals from the New York Central Zoo escape, unwittingly assisted by four absconding penguins, and find themselves in Madagascar.