When Gru, the world's most super-bad turned super-dad has been recruited by a team of officials to stop lethal muscle and a host of Gru's own, He has to fight back with new gadgetry, cars, and more minion madness.
The Madagascar animals fly back to New York City, but crash-land on an African nature reserve, where they meet others of their own kind, and Alex especially discovers his royal heritage as prince of a lion pride.
Manny, Sid, and Diego discover that the ice age is coming to an end, and join everybody for a journey to higher ground. On the trip, they discover that Manny, in fact, is not the last of the woolly mammoths.
After he is fired from the Anti-Villain League for failing to take down the latest bad guy to threaten humanity, Gru finds himself in the midst of a major identity crisis. But when a mysterious stranger shows up to inform Gru that he has a long-lost twin brother-a brother who desperately wishes to follow in his twin's despicable footsteps-one former super-villain will rediscover just how good it feels to be bad.