In a special Mother's Day edition of E! True Hollywood Story, E! charts the trailblazing journey of Joan & Melissa Rivers, the powerhouse duo who became role models for mothers and daughters everywhere. Beneath the constant one-liners and wisecracks, this moving story of a consummate entertainer and her producing partner is a testament to talent and family. Giving first-hand accounts of their lives, Joan and Melissa Rivers sat down with E! to tell their fascinating story - from Joan's early beginnings, when she defied her family to pursue her dreams of entertainment, to the heartbreaking suicide of Joan's husband and Melissa's father that almost destroyed the women's relationship, to their fabulously successful lives today.