The Enterprise stumbles upon a Dyson sphere, with a ship crashed on the outer surface. An away team finds some systems still powered up and the chief engineer from the old Enterprise, Montgomery Scott, locked in the transporter cycle.
While the crew is experiencing some much needed down time, Worf and his son go on an Old West excursion in the holodeck, where they meet several versions of Data serving as the bad guy after an experiment in Engineering goes wrong.
When Geordi is assigned to survey a new mining development with Dr. Farallon, who invented an experimental mining technology, she uses a new sophisticated robot called an Exocomp to repair a power grid.
Picard is replaced as captain of the Enterprise so he, Lt. Worf and Dr. Crusher go on a top-secret mission into Cardassian space. Meanwhile, his replacement, Captain Jellico, meets his new command with some resistance from the crew.
Murderous intrigue abounds for the Enterprise when one of the crew aboard a subspace station is believed dead, and suspected to have taken part in it until the Klingons show up with the young lieutenant, to Geordi's taste.
Aboard a shuttlecraft and en route back to the Enterprise, Picard, LaForge, Data, and Troi find themselves trapped in a time trap of some sort, where the Enterprise is being taken over by Romulans, and the ship, itself, on a course of destruction.
After an encounter with the Borg, Data feels his first emotion when he gets angry with the Borg. Data then tries to find ways to recreate the situation in order to feel emotions again, whilst the Enterprise investigates the Borg activity, and are bewildered as to why they feel emotions too.