New to England, Alexander Grayson hosts a lavish party. He becomes fixated on Mina, a beautiful young woman who looks like his dead former love. Newly engaged to Mina, Harker grapples with worries over providing Mina the life she deserves.
Grayson takes Lady Jayne as a lover even as he investigates her lethal connections to The Order of the Dragon. Grayson helps Mina face a critical challenge at medical school. Van Helsing continues his quest to develop a solar vaccine to enable Grayson to conquer sunlight. Jonathan Harker strikes a devil's bargain with Grayson which has unexpected consequences.
Grayson seeks to determine whether Lady Jayne is a vampire hunter for the Order of the Dragon. Lucy mends Mina's broken heart with absinthe, romance and a whirlwind tour of Bohemian London. Grayson's machinations claim the life of Lord Laurent and earn him a powerful new enemy.
Grayson ruthlessly sets out to compromise Lady Jayne, by winning her heart. Lucy's enthusiastic plans for Mina and Harker's engagement party conceals a broken heart. Grayson's unexpected reunion with a very old friend proves a growing complication to his and Van Helsing's mission of vengeance. Van Helsing achieves a breakthrough in his efforts to enable Grayson to conquer sunlight, only to suffer an unanticipated setback.
Mina Murray & Jonathan Harker's engagement ceremony takes place. Something pushes Mina to have second thoughts. Van Helsing continues his studies to make Alexander walk in daylight. Jane Wetherby is becoming suspicious about Mr. Grayson.
Browning sets out to prove once and for all that Grayson is the master vampire stalking London by hosting a board meeting outdoors in the sunlight. Grayson and Van Helsing seek to acquire the final component to make their wireless electricity technology a success to be witnessed. Meanwhile, Mina and Harker consummate their love - not knowing that Lord Davenport plots to destroy the object that Grayson loves most...Mina.