The Evil Queen sends Emma back to the Enchanted Forest and Regina must remind her of who she is. Belle and Rumplestiltskin try to figure out where their son is and David makes a wish that comes with a price.
Mary Margaret, Emma, Mulan and Aurora return to their camp only to find everyone dead. Back in Storybrooke, Regina begins to see a ghost from her past, as flashbacks show her training with Rumplestiltskin.
When a murder occurs in Storybrooke on a full moon, Spencer tries to bring the town against David by implicating Ruby, as flashbacks show Red Riding Hood find a pack of wolves who adopt her into their family.
Cora does everything in her power to steal the compass and find a way to Storybrooke, while after Aurora is kidnapped, and to protect Henry from further danger, Mary Margaret and David try to communicate with each other in the Netherworld.
As Hook and Cora arrive in Storybrooke, Regina struggles to gain the trust of the town, as flashbacks show Snow and Charming disagree over how to handle Regina now that they have taken over the kingdom.
Emma is shocked when she learns the true identity of Gold's son, as flashbacks show when Rumplestiltskin was called to war. Back in Storybrooke, Regina, Cora and Hook search for Rumplestiltskin's dagger.
Mary Margaret is tempted by dark magic and David, Emma and Neal attempt to protect Gold from Regina and Cora, as flashbacks show a young Cora seek revenge against the royal family after they degrade her in public.