Nick and Jess run off to Mexico to give their relationship a chance to grow without the everyday problems of Schmidt and Winston. Winston finds out he is color blind and cannot do puzzles. Schmidt makes a commitment to Cece and Elizabeth.
When Jess desperately tries to fit in with a clique of "mean girl" teachers at her new school, she turns to Nick for advice on how to deal with them. Meanwhile, an office party is anything but fun for Schmidt, and Winston plots revenge while cat-sitting for Daisy, whom he suspects is cheating on him.
Thinking Schmidt has chosen to be with Cece, Jess and Nick invite him to go on a double date with them. Nick is caught between loyalty to his friend and loyalty to his girlfriend when he finds out a shocking secret. Meanwhile, Winston promises to get the foursome a table at an ultra-exclusive restaurant that doesn't take reservations.
Now that Schmidt lost two girlfriends, he does not like seeing Nick and Jess so happy, so he tries to break them up. Winston realizes his cat has to be neutered, so he and the cat go out for one final festive evening before the operation.
Schmidt worries about being a good person. Nick inherits money from his father and starts a spending spree instead of paying his bills. Jess finds his box of bills and "stuff", and tries to help Nick by cleaning it out.
The guy's former roommate Coach is back in town and wants to party hearty with the guys as if time has not passed. Jess is upset that Nick is going to a strip club with Coach, so she and Cece have a girl's night out.
Coach tells Nick to give him a day to help change Nick's life with diet and exercise. Jess tries to get a program at school for conservation but the principal is against it. Menus for the Chinese restaurant keep getting delivered.
Jess is tasked with keeping a frantic Schmidt occupied inside the loft, as Coach and Cece go out on a first date. Meanwhile, Nick and Winston meet a frisky bus driver as they search for a missing Ferguson.
Coach accuses Nick of going soft since he started dating Jess. Nick decides to show his manliness by having everyone go camping for Thanksgiving, including hunting and foraging. Winston & Cece bond over not liking camping.
Jess wants to be Coach's friend. Nothing has worked so far until she realizes that her only way in is through his obsession with the Pistons. Nick is unhappy with Jess' new friendship as he is a Bull's fan.
Jess is ready to paint the town purple when a chance encounter finds her and Cece invited to a once-in-a-lifetime mansion party thrown by music legend Prince, guest-starring as himself. Not wanting to miss out on the fun, Nick, Schmidt, Winston and Coach are determined to crash the festivities, building toward an unforgettable ending. The special episode also features cameo appearances by Victoria Secrets' supermodels Alessandra Ambrosio and Ana Beatriz Barros and Los Angeles Dodger and Cy Young Award winner Clayton Kershaw.
When Nick has an awkward run-in with his ex, Caroline, Jessica tries to convince him he can still be friends with her, by introducing him to her former boyfriend, Berkley. Meanwhile, Coach uses Schmidt's loft to impress a date, but things turn farcically disastrous when Schmidt arrives home with his own potential bedmate.
Jess' sister visits and Jess does not want to introduce Abby to her friends because Abby has a way to ruin everything. Schmidt asks Nick to attend a Bar Mitzvah with him. Winston and Birdie have dinner with Coach and Cece.