Richard returns to talk about his horrifying 288 mph (460 km/h) crash in the Vampire jet-car that nearly cost him his life. The team take the Jaguar XKR through its paces, and set off on a challenge to prove that a 1.5 mile (2.4 kilometre) stretch of road can be resurfaced in 24 hours instead of the five days predicted by the local council.
James goes to Volkswagen's test track to drive the Bugatti Veyron at 253 mph (406 km/h), faster than any Formula One car. Hugh Grant is the Star In A Reasonably Priced Car. Jeremy, Richard and James drive the latest Audi TT, Alfa Romeo Brera and Mazda RX8 coupés to Scotland, where they each have two minutes to persuade the Scottish National Gallery Of Modern Art's experts that their car is a work of art.
Jeremy, Richard and James head to the United States, where they each have to buy a car in Miami for less than US$1000 and drive it across four states to New Orleans. One of the challenges they set along the way leads to the locals chasing them out of town.
The team put the latest Porsche 991 Turbo through its paces. Jeremy finds out why the Mercedes-Benz Brabus SL65 is the most powerful convertible yet. Richard, James and the British Amateur Rocket Society send a Reliant Robin into space on a rocket and turn it into a re-usable shuttle. Simon Pegg is the Star In A Reasonably Priced Car.
The team join forces with Railtrack to show what happens when a car that jumps the lights at a railway crossing is crashed into by a train. Jeremy takes the new Lamborghini Murcielago LP640 for a spin. Jeremy, Richard and James put some monster tractors through a series of challenges and use them to try growing their own biofuel crops. Kristin Scott Thomas is the Star In A Reasonably Priced Car.
Richard drives a Ford Shelby Mustang GT500. Jeremy, Richard and James each turn a working car into a stretch convertible and use it to take a high-profile celebrity client through London's narrow streets to the Brit Awards.
As part of Comic Relief 2007, the boys revived "Top Of The Pops" by having performances by Lethal Bizzle, Supergrass, Travis and Mcfly as well as integrating the Top Gear segment with music themes which includes news about upcoming music festivals etc.. In addition, the boys will perform for the very first time (Richard playing the bass guitar, Jeremy on drums and James on the keyboard) with singer Justin Hawkins.