Open with the gang practicing for the upcoming Glee Invitational show. The kids tell Will (Matthew Morrison
) they're worried they won't be able to succeed at upcoming competitions without Rachel (Lea Michele
). Finn (Cory Monteith
) tells Will he's not going to tell his family about Quinn (Dianna Agron
) being pregnant.
While at dinner with Terri (Jessalyn Gilsig
), Will meets a 24-year old still competing in a nearby high school's glee club.
Will told Emma (Jayma Mays
) about Quinn's pregnancy. Without revealing her knowledge of Quinn's situation, Emma takes a special in interest in Finn's future and suggests he try for a music scholarship. Performing well in the sectional and regional competitions may help drum up college interest, she tells him, and she suggests trying to bring Rachel back to Glee.
Finn finds Rachel working on publicity for the musical. He flirts openly with her and offers to help her run lines.
Will has Emma look up the file of April Rhodes (Kristin Chenoweth
), a classmate of his who was the biggest star of the McKinley High Glee Club and Will's first crush. He sees on her transcripts she came several credits shy of graduating.
Will finds April online, and she has him meet her at a luxury home. After pretending the mansion in hers, April is outed by a real estate agent as a squatter. Outside April tells Will she went to New York with her high school sweetheart and never ended up making it big. She agrees to come back to McKinley, take a few courses to graduate and join Glee again.
The Glee kids aren't convinced April can be as good as Rachel. To prove her skills, April performs "Maybe This Time," with Rachel joining in towards the end from the auditorium. April's so good, Kurt (Chris Colfer
Still, Will doesn't think the kids in Glee have accepted April socially. He suggests she try and win them over. This involves April introducing Kurt to booze, teaching Mercedes (Amber Riley
) and Tina (Jenna Ushkowitz
) how to shoplift and flirting mercilessly (and showering) with the football guys.
Sandy (Stephen Tobolowsky
) isn't happy with Rachel's work in Cabaret, primarily because he wants to install himself in.
Finn works with Rachel on her lines. He suggests they go bowling together to relieve some stress. Rachel and Finn are shocked when Will comes in to teach April "Don't Stop Believing," which had been Rachel's song.
A drunken Kurt throws up on Emma in the hallway. After several decontamination showers, Emma tells Will that April is a bad influence.
After Sandy yells at Rachel once again, she runs to the bathroom to cry. There she bumps into April, and the two have an argument. April promises Rachel she is never going to relinquish her renewed role as school star.
Finn and Rachel are at the bowling alley, as are Will and April. Will expresses his concern April is a poor influence, and she tells him she will go back on the wagon. He then confesses his dream was for them to perform together. They do a quick duet of Heart's "Alone." Finn tells Rachel everybody misses her in Glee. They eventually end up kissing, and he convinces her to quit the musical and return to Glee.
Puck (Mark Salling
) tells everyone in Glee that Quinn's strange behavior (puking, etc.) is the result of a "bun in the oven" and Finn is the father (even though Puck is). Just then Rachel walks in to rejoin Glee. Kurt tells her about Finn and Quinn's pregnancy, and Rachel is stunned.
After bringing Emma an application for a music scholarship, Finn is confronted by Rachel in the hallway with a satisfying slap. She is upset he didn't tell her about Quinn and says now she will never return to Glee. Sue (Jane Lynch
) gives Rachel complete creative control over Cabaret.
April shows up drunk for the Invitational performance. Emma thinks Will shouldn't let her go on. He decides yanking her would only be punishing the other kids. Glee goes on, with April doing a rousing rendition of Carrie Underwood's "Last Name." Rachel watches from the back, obviously upset.
After the first number, Will tells April he can't let her go back on stage. April agrees, telling Will the kids are lucky they have him to prevent their lives from turning out the way hers did. Needing a new lead, Rachel steps in -- sucking up her pride to be the "understudy" and helps Glee knock "Somebody to Love" out of the park in the second act.