Rachel proposes an idea to Mr. Shuester in order to improve the club's chance at the invitational. Meanwhile, Sue conjures up a plan (with the unintentional help of Sam) to destroy the Glee club once again.
Will thinks the glee club desperately needs more of an edge, so he brings in his former classmate, April Rhodes, to spice things up. Will's plan seems to be going smoothly until he realizes he may be losing a key member of the club.
Will challenges the kids to a healthy dose of competition with boys against girls in a mash-up showdown. Meanwhile, Terri takes a job as the school nurse, despite having no medical experience, to keep an eye on Will.
Inspired the by the students, Will composes a mash-up for a very special occasion. Meanwhile, in a shocking reversal of roles, McKinley High's cool kids are dethroned in an "icy" fashion, and Sue shows off her softer side.