Spoiled by their upbringing with no idea what wild life is really like, four animals from New York Central Zoo escape, unwittingly assisted by four absconding penguins, and find themselves in Madagascar, among a bunch of merry lemurs
Manny, Sid, and Diego discover that the Ice Age is coming to an end, and join everybody for a journey to higher ground. On the trip, they discover that Manny, in fact, is not the last of the wooly mammoths.
Ben Stiller returns as night watchman Larry Daily, now a successful business man, who gets back to the museum just in time to find that he needs to get his friends out of trouble. This new installment takes us to the Smithsonian, and introduces us to new characters, such as Amelia Earhart, General Custer, and many more!