Reed and Joel bury the hatchet to face a new tornado season, but armor offers little protection when the roof blows off the new Dominator. Sean mixes up his crew and unleashes tension in the path of a west Texas twister.
The deadliest tornado of the season tears a hundred mile path across Mississippi, destroying 600 homes and the town of Yazoo City. For Reed, Joel and Chris, an epic chase turns into a race to save lives, and changes their own lives forever.
After the worst tornado in his life, Reed pursues a new force of nature in TV weather woman Ginger Zee. But chasing more than weather leaves teammates out in the cold. As twisters rip through the Ozarks, one long-time chaser calls it quits.
Sean's team struggles with the death of a teammate, and heightened emotions lead to questionable driving. Reed arms the Dominator with probe cannons, while Tim smokes out a South Dakota monster that just keeps growing.
Storm and chasers collide in Minnesota for a final showdown. As a ferocious EF-4 twister targets the town of Wadena, one team finds themselves in the wrong place at the right time, and Sean's trial by fire becomes a race to save others.