Betty fires on all cylinders to get a big scoop with Gucci's new designer, Evan York, with the help of Hilda's connected customer, Sammy. But Marc plots to sabotage her, enlisting a clueless Amanda and Matt in his efforts, culminating in a showdown at the restaurant where Ignacio works. Meanwhile Daniel begins to attend a bereavement group, where he meets Natalie, a sexy girl who takes him out for an interesting night on the town, and Wilhelmina flies to Bermuda when she learns Connor may be there
Betty is overwhelmed with her editorial work, so Wilhemina assigns Marc to be Betty's temporary assistant. Betty's story idea to cover the worst jobs in New York goes south when she has to dress up as a hot dog and do a Bollywood dance number. Meanwhile, Marc frets that he'll never be taken seriously by his idol at Vogue magazine, Hilda gets Archie's campaign in trouble, Daniel hires a gorgeous but inept assistant, and Natalie introduces Daniel to the Community -- a group whose charismatic leader zeroes in on Daniel.
Matt asks someone else to be his "plus one" at Wilhemina's fund-raiser for Tibetan orphans, and Daniel hires his new BFF Natalie as his assistant, making Betty feel obsolete. Amanda keeps a secret from Betty, and Daniel discovers that his mother, Claire, has history with Cal Hartley. Meanwhile we learn the truth behind Nico's dire straits.
Marc helps Justin to fit in high school by telling him to become friends with some mean girls. Daniel is lured into a trap by Bennett. Wilhemina tries to come up with money to help Nico out of trouble.
Daniel lets the leaders from the Community of the Phoenix call the shots at Mode, and Bennett's promises to reunite Daniel with his dead wife prompt Betty to action. She schemes with Claire on breaking through to him with an ingenious plan and finds unlikely allies in Matt and Amanda. Meanwhile, Wilhemina has let herself go, wearing sweats and garden clogs in public, prompting a tabloid feeding frenzy and spurring Marc to dig deeper into Nico's mystery.
In the steamy Bahamas, the love triangle between Betty, Matt and Amanda explodes, and romantic pairings abound. Things should be dreamy at the Atlantis resort, where even Shakira is staying, but Betty has a nightmarish photo shoot after Willie learns her nemesis, Penelope Graybridge, snagged a coveted job, and, even more importantly, discovers Connor is very much alive. Marc frets that there is no one on the island for him, but then meets the uber-hot Troy, and back home, Hilda draws closer to her old love, Bobby,
Against the backdrop of Ignacio's multi-culti holiday dinner that he prepares for his new flame Jean, who is Jewish, Betty and Hilda have pregnancy scares. Worse, they purchase their Be-Shure instant pregnancy tests from the local pharmacy -- where Jean works and Ignacio is shopping. Meanwhile, Hartley installs Willie's replacement, a clueless Hollywood mogul, Denise Ludwig, whose claim to fame is a tacky movie, but Marc has a plan to get Willie back -- if only Daniel will agree. And Claire gets Amanda's help and tracks down Tyler, the son she long ago gave up for ...
When Betty becomes concerned that she and Matt are spending too much time together, she tries to give him something else to be passionate about -- a surprise gallery showing of his art work -- with unintended results. Meanwhile, Marc is horrified that his one-night stand, Troy, is falling for him, since Marc was Troy's "first," Hilda grapples with whether or not to tell Bobby that he's the father of her unborn child, and Cal Hartley hires a Hollywood powerhouse, Denise Ludwig, to replace Wilhemina.
A battered but unbowed Betty recounts her worst week ever to Mr. Z as the show flashes back to her horrible last few days. Frustrated over the frivolous assignments Wilhemina gives her at Mode, and inspired by Audrey Hepburn's memoir, Betty creates her own blog about amazing charitable ventures that inspire her. However, as her hobby enriches her, her job is nearly killing her, since Wilhemina tortures Betty with beauty experiments for future stories. Meanwhile, Daniel fears Marc is sabotaging him -- and he is; Bobby treats a hormonal Hilda to some pampering during "...
Betty tries to be neighborly by planning a karaoke party in her apartment building, but before the big day, her building is robbed, and it's partly her fault. Marc is distraught since he lost something irreplaceable in the theft. Meanwhile, Daniel and Wilhelmina are back to being at each other's throats, so Amanda tricks them into attending couples counseling with Dr. Sparks, and Ignacio refuses to accept Bobby in Hilda's life. Then a citywide blackout ups the ante for everyone.
Wilhemina is furious when she discovers she has a drag queen impersonator - Wilhediva Hater. Meanwhile Betty receives an award for her blog, which causes tension between her and Daniel. When she vents online, her fans in blogosphere escalate the feud, and the press hounds Daniel. Amanda rehires Helen, but Willie orders Amanda to fire her -- which Amanda just can't bring herself to do. And Marc runs his own photo shoot with a famous director and actress.
Wilhemina marginalizes Betty during Fashion Week but Daniel intercedes, and she's assigned to cover a low-priority show. There, she discovers a new designer, Marisa, whom she promotes as one of the 10 Designers to Watch, but Marc steals all her thunder. Meanwhile, Amanda is impressed by Helen's sewing skills and teams up with her to start a new line, but cannot convince Marc or Betty to lobby Wilhemina on her behalf. Claire's son Tyler comes to New York to find her, and Daniel gets seduced into modeling. The fashion and the feelings are smoking hot, but the real ...
To speed along the investigation into her house fire, Betty is pressured to date an annoying fireman. Wilhemina tries to prove to Don, an old flame from her past, that she's just "Wanda from the block" instead of an uber-bitch, and Amanda draws closer to Tyler, convincing Claire to hire him as an in-house model at Mode. Meanwhile, Hilda meets Bobby's parents for the first time.
Justin loves his acting class, helmed by the eccentric Lena Korvinka, and finds himself in an unexpected triangle with his new pals, Austin and Lily. When Betty drops by class, she's smitten with a cool young playwright, Zachary, whom she will profile for Mode. But she challenges her own ethics when she starts dating him. And Wilhemina is hospitalized with a perforated ulcer, and can't help her type-A personality even when bedridden, but then her roommate, Roberta, prompts an epiphany.
Betty is thrilled to get her braces off, courtesy of her chatty new orthodontist, Dr. Frankel. But before this can happen, she gets knocked unconscious at a photo shoot of the "Million Dollar Bra" at the Guggenheim, spinning her into a dream sequence. An angelic Dr. Frankel shows her how different her life would have been had she never worn braces. She's the "pretty sister," while Hilda is homely, Ignacio is a compulsive gambler, Marc is a doting dad and Amanda is married to Daniel. But glam Betty is also mean - she's in cahoots with Wilhemina to terrify and overpower...
Betty gets an assignment to cover London Fashion Week, accompanied by Amanda, and she brings Hilda along as a bachelorette party treat. She reconnects w/Christina and also runs into Gio unexpectedly. Willie joins AA to get closer to Tyler and to double-cross Claire and Daniel, and Bobby can't figure out why Justin is spending all his time with Austin -- until he can.
As Hilda's wedding day approaches, Betty asks Daniel to be her plus-one, Ignacio invites Elena, and Amanda is set on fixing up Marc with her new client, soap opera star Spencer Cannon. Meanwhile, Justin weighs whether his own plus-one could be Austin, Betty considers unexpected offers in her personal and professional life, and Wilhelmina finds a way to fight the Meade empire by using Tyler as her pawn.
It's the end of an era, but the beginning, Betty hopes, of something wonderful. In the series finale, which also features appearances by familiar faces from the past, Betty finds it hard to tell Daniel important news about her future. When she does, his reaction surprises her, but not Claire. Justin and Bobby enthusiastically consider taking advantage of a softening real estate market by apartment-hunting in Manhattan, but Hilda's resisting even the most amazing places. Amanda grapples with the death of her dog, Halston, and Marc wonders if he'll ever have a ...