The station hires financial analyst Julia Wilcox, an abrasive and sharp-tongued woman with no time for anyone. During a particularly vicious argument with her, Frasier becomes aroused and makes an advance, leading to a mandatory sexual harassment training session for the entire staff.
Now that Kenny's divorce is final, the realization that his wife is never coming back hits him hard. Frasier suggests he see a psychiatrist, and Kenny asks him to be his "shrink". Frasier reluctantly agrees, but then is energized by the thrill of deeply analyzing a patient again. Kenny, on the other hand, doesn't care for the long sessions. After going out for a drink with Martin, he realizes that he just wants to have some fun and ends his sessions with Frasier, much to his disappointment.
Frasier and Niles acquire an inside source of top-quality caviar, and become "dealers" to the rest of Seattle's elite, dramatically improving their status. Martin wrestles with a bank's bureaucratic procedures, trying to return a trivial overpayment on his account.
After the KACL staff participate in a low-income house-building project, Frasier becomes a nuisance to the family, offering incessant advice for its decor. Martin and Eddie score a success with a safety assembly at an elementary school, but Niles must fill in at the next one when Martin gets sick.
Julia breaks off her affair with Avery, Frasier's accountant. She breaks into his office to steal back some of her work. Frasier follows her, and as he constantly urges her to do the right thing, she succumbs to a sudden attraction and kisses him. Roz is offered a plum position at another radio station. Niles develops a passion for target shooting.
Unsettled by Frasier's burgeoning relationship with Julia, Roz reverses her decision to leave KACL, but issues an ultimatum to Frasier: dump Julia, or they are no longer friends. Frasier refuses, and Roz takes the new job after all. Niles and Daphne decide it is time to try and conceive, and work up the nerve to order Gertrude out of the house.
Roz is unsatisfied with her new job, and comes back to KACL, acting as though nothing has changed. Frasier believes her attachment is based on deep attraction to him stemming from their one-night stand. Roz disabuses him of this notion, and they go back to being friends. Recalling that he once donated sperm, Niles returns to the bank to see whether it was ever used, and is told that he suffers from low motility. He tries a variety of weird medical remedies.
After breaking up with Julia, Frasier lapses into his usual self-pitying funk. Niles reminds him that he has a fear of commitment that leads him to sabotage all his relationships with women. Frasier typically overdoes things, committing to commitment with Julia, even during an evening with his family when she rubs everyone the wrong way, and steps on Niles and Daphne's surprise announcement that she is pregnant.
Frasier decides to resume private practice. While shopping for a suitable couch, he and Niles catch sight of Ronee Lawrence, their old babysitter. Frasier invites her for a drink, but it quickly becomes apparent that she is interested in Martin.
Roz sets Frasier up with her friend, Ann, advising Frasier that he should keep his dating skills in shape, even if it means going out with someone he's not interested in. Frasier agrees, but their first date is a night in Hell, made worse by the fact that Kenny is dining in the same restaurant with his attractive and eminently suitable cousin. Niles and Daphne have a fastidious cleaning lady whom Niles adores, but Daphne believes is stealing.
Frasier inadvertently eavesdrops on Ronee making a date with another man. Martin takes the news badly, accusing him of spying and inventing a fictitious girlfriend to spark Ronee's jealousy. Niles tries to win the attention of a reclusive nursery painter.
Maris meets with Niles, seeking protection from her violent new boyfriend. Because of Daphne's mood swings, Niles tries to keep his meeting secret, which fails when the boyfriend shows up at a fake baby shower and punches Frasier. Frasier's first day back in private practice goes less smoothly than he hoped.
Maris is arrested for the murder of her boyfriend. The press goes wild when they find out that Niles inadvertently loaned her the crossbow that was used as the murder weapon. Niles slowly goes to pieces under the unrelenting media scrutiny and Daphne's anger.
Lilith is in town for a psychiatric conference, and Frasier reluctantly assumes his obligation to meet with her. Before he does so, an interested colleague of Lilith's sets him up on a blind date with her, not knowing who he really is. Martin's pistol accidentally discharges inside the apartment, creating a trail of destruction that he, Daphne and Niles try to hide from Frasier.
For once, Niles is the famous one, as media attention to Maris's case makes him and Daphne the center of Seattle's attention. Frasier is left to contemplate his falling ratings and his dating woes. In a funk, he agrees to emcee the SeaBea Awards and give an interview to a newspaper reporter, neither of which goes well. Roz seethes during a visit from her "perfect" sister, Denise.
Frederick visits Frasier in Seattle, and the family is shocked to see him going through a "Goth" phase. The family talks about the rebellious phase of a child's life, and Niles is stung to hear Martin say that he was always one of the good kids. He decides to rebel by consuming a pot brownie supplied by Roz, but hijinks ensue when Martin eats it.